Morphogenesis – “The - 麻省理工学院人工智能研究专业

Morphogenesis – “The
Riddles of Form” in
Twenty-First Century
科学

Marco Tamborini
Department of Philosophy
Technische Universität
Darmstadt, 德国

Over the past decades, the notions of organic form and morphology—a
scientiﬁc ﬁeld historically associated with the eighteenth century polymath
Johann Wolfgang von Goethe (1749–1832)—have stealthy re-assumed a
central role in various scientiﬁc disciplines. Although the study of organic
form was apparently excluded from the main stage of evolutionary theory
and biological sciences during the second half of the twentieth century, 自从
morphology was considered as a descriptive and ancillary science unable to
contribute to the neo-Darwinian synthesis of evolution1, morphological con-
cepts and approaches have now been re-brought onto the central stage of
mainstream science.2

实际上, several interdisciplinary Clusters of Excellence3 have been for
instance established in Germany to investigate the enigma and power of

See Ghiselin 1980, 2006; Mayr 1980. For an alternative view see Hopwood 2003;
Nyhart 1995; Love 2006; Tamborini 2020c, under contract; Levit, Hossfeld, and Olsson 2014.
2. On the philosophical question of using old issues to pose new questions see, 为了

例子, Gutmann and Tamborini 2020; Baedke 2019.

3. As part of the Excellence Initiative the German Universities Excellence Initiative
supports pioneering research. Through a massive ﬁnancial support, the Clusters of Excel-
lence aim at reaching outstanding research and obtaining international recognition and
visibility. As the German Research Foundation DFG reported on its website, “[C]lusters
of excellence enable German university locations to establish internationally visible, com-
petitive research and training facilities, thereby enhancing scientiﬁc networking and coop-
eration among the participating institutions.” For morphological-oriented clusters see, 为了
实例, the Clusters of Excellence “Integrative Computational Design and Construction
for Architecture” led by Achim Menges at the University of Stuttgart and the Cluster of
Excellence “Matters of Activity. Image Space Material” lead by Wolfgang Schäffner at
Humboldt University in Berlin and “Living, Adaptive, and Energy-autonomous Materials
Systems” led by Jürgen Rühe at the University of Freiburg.

科学观点 2021, 卷. 29, 不. 5
© 2021 由麻省理工学院

https://doi.org/10.1162/posc_e_00383

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560

介绍 – Morphogenesis

form transformations. These clusters recall and expand Goethe’s and
Romantic deﬁnition of form, this meant as a complex and ever-changing
现象. Within biological disciplines, new research programs have
been put forward to explicitly investigate what has been disqualiﬁed in
biology over the past decades: the very concept of organic form. 例如,
according to Austrian evolutionary biologist Gerhard Müller, “Evolutionary
developmental biology (evo–devo) emerged as a distinct ﬁeld of research
in the early 1980s to address the profound neglect of development in the
standard modern synthesis framework of evolutionary theory, a deﬁciency
that had caused difﬁculties in explaining the origins of organismal form
in mechanistic terms (穆勒 2007, p. 943). 此外, synthetic biolo-
gists and nanoscientists are now producing and manipulating a multitude
of forms to design and control their possible genesis (例如, Gramelsberger
2020).

In addition to the renaissance of morphological studies in evolutionary
生物学, the notion of organic form has deeply inﬂuenced twentieth- 和
twenty-ﬁrst century architectural and computational design. In the mid-
1970s, German biologist Johann-Gerhard Helmcke (1908–1993), WHO
collaborated with German architect Frei Otto (1925–2015) in establishing
two Collaborative Research Centers4 (Sonderforschungsbereiche – short SFB),
写了: “I wondered if the architects could recognize the beauty of biological
objects and then ﬁnally build something more aesthetically, and if engineers
could understand the many innumerable, biological forms of evolution of
constructions in order to learn from them and to build better (也许
also more economically)” (Helmcke5).

Helmcke’s hope was rooted in a deep history of interaction and entangle-
ment between architecture and organic morphology (例如, Steadman 2008).
This collaboration found partial international recognition with the estab-
lishment of two Collaborative Research Centers in West Germany between
early 1970 and late 1980. 在 1970, the so-called SFB 64 on “Weitgespannte
Flächentragwerke” [Wide Span Surface Structures] was established in
Stuttgart. This developed into the SFB 230 on “Natürliche Konstruktionen”
[natural constructions] starting from 1984. The peculiarity of these Col-
laborative Research Centers was that biologists, engineers, architects, artists
worked together to investigate form’s intrinsic dynamics (see Tamborini
2020A). The intertwinement between architecture and biology, 或者更确切地说

In the German system, Collaborative Research Centers “are long-term university-
based research institutions, established for up to 12 年, in which researchers work to-
gether within a multidisciplinary research programme” (DFG).

5. Nachlaß 135. Staatsbibliothek zu Berlin – Stiftung Preußischer Kultur-besitz.

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科学观点

561

between a technical and a biological study of form transformation and
manufacturing, reached its heyday in the development of architectural
and computational design during the 2010s.

Commenting on these recent developments in an article emblematically
entitled Digital Morphogenesis, British architect and theorist Neil Leach noted
that “architecture [……] is no longer so preoccupied with style and appear-
安斯. It is as though a new paradigm has emerged [……] the more contem-
porary architects operating within the new morphogenetic paradigm can be
seen more as the controllers of processes, who facilitate the emergence of
bottom-up form-ﬁnding processes that generate structural formations”
(Leach 2009, p. 34).

此外, alongside fostering the production of bio-inspired con-
structions, the biological notion of organic form is shaping the current
advancement of computational design. 今天, architects and designers are
proposing a shift from a modernist notion of form analysis, which empha-
sized the design of well-adapted forms, towards computational research,
which seeks to set the organizational rules responsible for form generation.
实际上, by means of computational tools, architects design the code whose
guides form development. Within this computational perspective, “form
[becomes] a subsidiary component of environment, and environment …
a complex web of inﬂuences” (Menges and Ahlquist 2011, p. 10). 这
移动, as architects Achim Menges and Sean Ahlquist noted, “represents
an accumulation of multilayered concepts ranging from system theory
and cybernetics, to morphogenesis and developmental biology” (Menges
and Ahlquist 2011, p. 10; see also Greg Lynn’s works, 例如, 林恩
1999).

What is the implication of this renewed interdisciplinary interest in
form transformation and production? Is it a mere return to a romantic-
holistic conception of nature, or rather does it foster a new standpoint
on nature according to which nature is meant as the product of engineer-
设计? What kind of knowledge and practical skills are associated
with such bio-technical understanding of organic forms? How did digital
technologies and different engineering approaches to biology impact and
shape the classical morphological traditions? Succinctly, what is so enigmatic
in the concept of organic form?

This special issue investigates in-depth all these questions by present-
ing a more nuanced picture of the twentieth and twenty-ﬁrst century
study of form. It analyzes to what extent morphological knowledge pro-
duction is tied with and rooted in different technological settings and
broader philosophical frameworks. In all the cases mentioned above,
knowledge production and product design and manufacturing are deeply
rooted in romantic-holistic morphological concepts. These concepts were

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介绍 – Morphogenesis

re-formulated and expanded to emphasize the idea of a possible technical
or engineering control of form changes.6

Famously, Goethe deﬁned morphology as the science [Wissenschaft] 有能力的
to “recognize living formations as such, to grasp their externally visible,
tangible parts in relation to one another, to take these parts as indications
of what lies within and thus to acquire a degree of mastery over the whole
in intuition” (Goethe 1817 (2000), p. 56; translation in Steigerwald 2002,
p. 314). The German polymath meant though this discipline in a very
dynamic way. 他, 实际上, emphasized a strong linguistic and conceptual
difference between two different German expressions for form: “The German
has a word, Gestalt, for the complex existence of an actual being. He abstracts,
with this expression, from the moving and assumes a congruous whole to be
determined, completed, and ﬁxed in its character … But … we ﬁnd that
独立, 休息, or termination nowhere appear, but everything ﬂuc-
tuates rather in continuous motion. Our speech is therefore accustomed
to use the word Bildung pertaining to both what has been brought forth and
the process of bringing forth” (Goethe [1817] 2000, p. 55; translation in
Zammito 2017, p. 483).

By so doing, Goethe rejected a static form deﬁnition. He emphasized,
反过来, its dynamic and ever-changing status. 因此, the German
poet and polymath conceived morphology as morphogenesis. That means
as the study of the dynamics of form formation and change over time. As he
把它, morphology was “the theory of form [Gestalt], 形成 [Bildung]
and transformation [Umbildung] of organic bodies” (Goethe [1817] 2000,
p. 124, translation in Steigerwald 2002, p. 295).

Following Goethe’s ideas, at the end of the nineteenth century, morphol-
ogy was considered by most biologists as “the ﬁrst evolutionary science”
(Bowler 1996, p. 17) because of its pivotal role in analyzing and understand-
ing evolutionary changes thorough time (Rieppel 2016; 坦博里尼 2019).
During the twentieth century, pushed by morphology’s exclusion from the
so-called modern synthesis of evolution several morphologists became in-
creasingly less interested in bringing out historical and phylogenetic expla-
nations. They lost their interest in drawing phylogenetic trees or series of
形式, though still elevating Goethe as their patron saint7. In redeﬁning
and expanding their disciplinary space of knowledge, many morphologists
started adopting new technologies and developed so-called engineering
approaches to the study of evolution.

6. On the intersection between romantic conceptions of life and technology see also

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Esposito 2016; 彼得森 2016; Harrington 1999; Tamborini 2020b.

7. As Olivier Rieppel noted, they appreciated Goethe’s theory of morphology in its

original version as ahistorical investigation of form’s structures (Rieppel 2020).

科学观点

563

The Scottish polymath D’Arcy Thompson (1860–1948) epitomized this
engineering interest in the form study. He expanded the Goethean notion
of morphology and morphogenesis to both natural and organic phenome-
非: “The waves of the sea, the little ripples on the shore, the sweeping curve
of the sandy bay between the headlands, the outline of the hills, the shape of
the clouds, all these are so many riddles of form, so many problems of mor-
phology” (汤普森 [1917] 1942, p. 10). All these riddles of form could be
technically grasped. They were generated as “a quasi-mechanical effect on
Matter of the operation of chemico-physical forces” (汤普森 [1917]
1942, p. 81). 实际上, Thompson supported the idea that “Form [……] 的
any portion of matter, whether it be living or dead, and the changes of form
which are apparent in its movements and in its growth, may in all cases alike
be described as due to the action of force. 简而言之, the form of an object is a
‘diagram of forces’” (汤普森 [1917] 1942, p. 16).

The transition from a purely biological, and historical, study of form
towards a hybrid one, in which engineering and biological sciences worked
一起, culminated in the establishment of bionics and biomimetics as
autonomous scientiﬁc disciplines. These were in turn considered as essen-
tial components of the broader research agenda pursued by architects and
biologists—for instance, architect Frei Otto invited biologist Werner
Nachtigall, founder of the ﬁeld of bionics in Germany, to formally partic-
ipate in his SFBs. 此外, biologist Helmcke set up with German
engineering Heinrich Hertel, professor of Aeronautical Engineering at
the Technical University of Berlin, a “marriage between engineering and
biology” which led to the battle cry ‘TUB’ [short for: Technologie und
Biologie], technology and biology (例如, Hertel 1963; Tamborini 2020a).
By bringing together historians of science, philosophers of science, 设计
理论家, and cultural historians, the papers collected in this special issue
problematizes the encounter between the romantic tradition of form study,
the engineering approach to evolution, and to biological approach to form
design and manufacturing. The analysis of this intertwinement will help us
reconsider classical philosophical notions, such as representation, 模型,
机制, and organicism. 此外, it will shed light on the dynam-
ics of knowledge production in nature inspired sciences, like biomimetics,
synthetic biology, and computational architecture.

Alfred Nordmann opens the special issue with some general reﬂections
on what biotechnology. He takes up categories from aesthetics to investi-
gate to what extent synthetic biology is copying natural form. He argues
that synthetic biologists do not copy the organic world, but rather they
create its parody. In his essay, he develops this idea further presents “a vo-
cabulary for the parodistic qualities of Synthetic Biology, genome editing,
or other bioengineering practices.”

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介绍 – Morphogenesis

In his paper, Ulrich Krohs aims to better understand the epistemology
of biomimetics. He ﬁrst reconstructs the biomimetic copying practice. 后
having proposed a four-step model to understand how biomimetic copying
activity does work, he investigates the differences between biological robust-
内斯, system’s ability to “maintain its functionality across a wide range of
operational conditions”, and robustness in technical and artefacts. This dis-
tinction enables him to examine the possibility of transferring robustness
from a biological to a technical system. Working on the transferability pro-
cess in synthetic biology, Krohs points out the differences between morpho-
genesis in technical and biological systems. He concludes by asserting that
“transferability between biological and technological systems is by and large
a one-way system”. That means that there is a sort of mimetic asymmetry.
The transfer “of the implementation of the function from the technical to the
biological realm” nicely works, but the transmission of “the means to achieve
robustness” miserably fails.

Carolin Höﬂer examines recent changes in the concept of organic forms
in architecture. 尤其, she focuses on the structural architecture of
the 1960s and the computer-based architectural systems since the 1990s.
She shows that from the mid-20th century onward, a renewal of architec-
tural design practice was sought through the reformulation of classical
morphological questions at the intersection of biological and cybernetic
话语.

George Toepfer argues that morphology can fulﬁll an important explan-
atory function and it constitutes the fundamental explanatory level of biol-
ogy as a distinct science. He further argues that the forms of organisms and
their parts provide the only speciﬁcally biological causal factors. After a clas-
siﬁcation of four types of morphology, he develops the thesis that organic
“forms can be viewed as forces or causes. They are the only other forces of
life beyond the forces of physics.”

Marco Tamborini situates his contribution at the intersection between
Krohs’ and Toepfer’s discussion of biomimetics and Höﬂer’s analysis of
型号. He examines the features of twenty-ﬁrst-century robots-inspired
morphology. After having distinguished biomimetics, or nature-inspired
工程, from robotics-inspired morphology as two complementary
学科, he investigates in-depth what role do robots play in the current
study of form. This brings him to focus on the features of what he named
“the recent material turn in morphology”. This turn is marked by the hybrid
co-existence of technologies, 机器人技术, and living organisms in studying
form’s structures and changes.

Mathias Gutmann concludes the special issue with a broader methodo-
logical and philosophical reﬂections on the notion of form. He develops a
conceptual framework for the notion of “form” useful to understand to

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科学观点

565

what extent this is still central in current biology. If form is meant as a meta
概念, argues Gutmann, we are able to better understand the practices,
程序, and argumentative framing that characterize twentieth- 和
twenty-ﬁrst-century biology.

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